Research
My primary research interest is in exploring philosophical issues in the measurement of animal welfare, and how this can apply to an empirically-informed animal ethics.
Future research interests include how to identify indicators of consciousness/sentience, how the evolution and mechanisms of sentience can inform welfare measurement, performing comparisons of sentience/welfare between different species, how to quantify the badness of death for animals and how much suffering is experienced by wild animals.
Future research interests include how to identify indicators of consciousness/sentience, how the evolution and mechanisms of sentience can inform welfare measurement, performing comparisons of sentience/welfare between different species, how to quantify the badness of death for animals and how much suffering is experienced by wild animals.
Publications
Books | Single-author papers | Coauthored papers | Coauthored commentaries and book reviews | Public engagement
Details of all publications can also be found on my Google Scholar profile
Books
Browning, H. & Veit, W. (2024). What Are Zoos For? Bristol: Bristol University Press.
https://bristoluniversitypress.co.uk/trade/what-are-zoos-for
Browning, H. & Veit, W. (2024). What Are Zoos For? Bristol: Bristol University Press.
https://bristoluniversitypress.co.uk/trade/what-are-zoos-for
Single-Authored Works
Browning, H. (2026). Mapping profiles of animal affect. Biology and Philosophy. 41(16): 1-23
https://doi.org/10.1007/s10539-026-10019-7 [Download]
Browning, H. (2023). Validating indicators of subjective animal welfare. Philosophy of Science. 90(5): 1255 - 1264
https://doi.org/10.1017/psa.2023.10 [Download preprint]
Browning, H. (2023). Welfare comparisons within and across species. Philosophical Studies. 180: 529-551
https://doi.org/10.1007/s11098-022-01907-1 [Download]
Browning, H. (2023). Improving welfare assessment in aquaculture. Frontiers in Veterinary Science. 10: 1060720
https://doi.org/10.3389/fvets.2023.1060720 [Download]
Browning, H. (2022). Assessing measures of animal welfare. Biology and Philosophy. 37(36): 1-24 https://doi.org/10.1007/s10539-022-09862-1 [Download preprint]
Browning, H. (2022). The measurability of subjective animal welfare. Journal of Consciousness Studies. 29(3-4): 150-179
https://doi.org/10.53765/20512201.29.3.150 [Download]
Browning, H. (2020). If I Could Talk to the Animals: Measuring Subjective Animal Welfare. PhD Thesis (Australian National University).
https://doi.org/10.25911/5f1572fb1b5be [Download]
Browning, H. (2020). The natural behavior debate: Two conceptions of animal welfare. Journal of Applied Animal Welfare Science. 23(3): 325-337
https://doi.org/10.1080/10888705.2019.1672552 [Download preprint]
Browning, H. (2019). What is good for an octopus? Animal Sentience. 26(7)
https://doi.org/10.51291/2377-7478.1476 [Download]
Browning, H. (2019). What should we do about sheep? The role of intelligence in welfare considerations. Animal Sentience. 25(23)
https://doi.org/10.51291/2377-7478.1460 [Download]
Browning, H. (2018). Won’t somebody please think of the mammoths? De-extinction and animal welfare. Journal of Agricultural and Environmental Ethics. 31(6): 785-803
https://doi.org/10.1007/s10806-018-9755-2 [Download preprint]
Browning, H. (2018). No room at the zoo: Management euthanasia and animal welfare. Journal of Agricultural and Environmental Ethics. 31(4): 483-498
https://doi.org/10.1007/s10806-018-9741-8 [Download preprint]
Browning, H. (2018). Book Review: Zoo Ethics. Quarterly Review of Biology. 93(2): 149
https://doi.org/10.1086/698059 [Download]
Browning, H. (2017). Anecdotes can be evidence too. Animal Sentience. 16(13)
https://doi.org/10.51291/2377-7478.1246 [Download]
Browning, H. (2026). Mapping profiles of animal affect. Biology and Philosophy. 41(16): 1-23
https://doi.org/10.1007/s10539-026-10019-7 [Download]
Browning, H. (2023). Validating indicators of subjective animal welfare. Philosophy of Science. 90(5): 1255 - 1264
https://doi.org/10.1017/psa.2023.10 [Download preprint]
Browning, H. (2023). Welfare comparisons within and across species. Philosophical Studies. 180: 529-551
https://doi.org/10.1007/s11098-022-01907-1 [Download]
Browning, H. (2023). Improving welfare assessment in aquaculture. Frontiers in Veterinary Science. 10: 1060720
https://doi.org/10.3389/fvets.2023.1060720 [Download]
Browning, H. (2022). Assessing measures of animal welfare. Biology and Philosophy. 37(36): 1-24 https://doi.org/10.1007/s10539-022-09862-1 [Download preprint]
Browning, H. (2022). The measurability of subjective animal welfare. Journal of Consciousness Studies. 29(3-4): 150-179
https://doi.org/10.53765/20512201.29.3.150 [Download]
Browning, H. (2020). If I Could Talk to the Animals: Measuring Subjective Animal Welfare. PhD Thesis (Australian National University).
https://doi.org/10.25911/5f1572fb1b5be [Download]
Browning, H. (2020). The natural behavior debate: Two conceptions of animal welfare. Journal of Applied Animal Welfare Science. 23(3): 325-337
https://doi.org/10.1080/10888705.2019.1672552 [Download preprint]
Browning, H. (2019). What is good for an octopus? Animal Sentience. 26(7)
https://doi.org/10.51291/2377-7478.1476 [Download]
Browning, H. (2019). What should we do about sheep? The role of intelligence in welfare considerations. Animal Sentience. 25(23)
https://doi.org/10.51291/2377-7478.1460 [Download]
Browning, H. (2018). Won’t somebody please think of the mammoths? De-extinction and animal welfare. Journal of Agricultural and Environmental Ethics. 31(6): 785-803
https://doi.org/10.1007/s10806-018-9755-2 [Download preprint]
Browning, H. (2018). No room at the zoo: Management euthanasia and animal welfare. Journal of Agricultural and Environmental Ethics. 31(4): 483-498
https://doi.org/10.1007/s10806-018-9741-8 [Download preprint]
Browning, H. (2018). Book Review: Zoo Ethics. Quarterly Review of Biology. 93(2): 149
https://doi.org/10.1086/698059 [Download]
Browning, H. (2017). Anecdotes can be evidence too. Animal Sentience. 16(13)
https://doi.org/10.51291/2377-7478.1246 [Download]
Coauthored Articles and Book Chapters
Veit, W. & Browning, H. (forthcoming). Do animals understand death, and does it matter? In The Palgrave Handbook on the Philosophy of Death.
Browning, H. & Veit, W. (forthcoming). From animal to AI welfare. In Z. Radman (ed.). HumANimal: The “Animal Turn” Meets Artificial Intelligence. Palgrave MacMillan.
Browning, H. & Veit, W. (forthcoming). Does the welfare of wildlife matter? In A. Simmons (ed.). The Welfare of Wildlife.
Veit, W. & Browning, H. (forthcoming). Animal desire. In A. Gregory (ed.). The Routledge Handbook on the Philosophy of Desire.
Browning. H. & Powell, D. (2026). Navigating welfare contradictions in zoo-supported conservation activities. Zoo Biology: e70100
https://doi.org/10.1002/zoo.70100 [Download]
Webb, L.E., Bolhuis, J.E., Jurkovich, V., Ipema, A.F., Chincarini, M., Favreau-Peigné, A., Forkman, B., Clouard, C., Tuyttens, F., Browning, H., Menchetti, L., Bak Jensen, M., Elmaz, Ö ., Šárová, R., Głogowski, R., Duepjan, S., Celozzi, S., Veit, W., Tallet, C. (2026). Balancing the animal affective scales - Can the methodology of human affect balance be translated to other sentient species? Neuroscience & Biobehavioral Reviews. 106914
https://doi.org/10.1016/j.neubiorev.2026.106914
Elliott, V., Beausoleil, N., Browning, H., Díez-León, M., Goddard, P., Graham, M., Ramp, D., Sebo, J., Soryl, A. & Zemanova, M.A. (2026). The emerging field of wild animal welfare science. Trends in Ecology & Evolution. Online first.
https://doi.org/10.1016/j.tree.2026.06.003 [Download]
Keens Caballero, H., Browning, H., Lambton, S., Maye, D., & Roe, E. (2026). Resilience ontologies in veterinary science: How they shape the way we address resilience. Veterinary Sciences, 13(5): 471.
https://doi.org/10.3390/vetsci13050471 [Download]
Veit, W. & Browning, H. (2026). Two kinds of conceptual engineering. Croatian Journal of Philosophy. 26(76): 49-70.
https://doi.org/10.52685/cjp.26.76.4 [Download]
Schnell, A.K., Browning, H., Crump, A., Burn, C.C. & Birch, J. (2026). Sentience in cephalopod molluscs: An updated assessment. Biological Reviews. 101: 1311-1333
https://doi.org/10.1002/brv.70125
Ishida, S., Kagan, B.J., Kataoka, M., et al. (2026). Ethics and regulation of human brain organoid research: Recommendations from the Asia Pacific Neuroethics Working Group. Asian Bioethics Review. Online first.
https://doi.org/10.1007/s41649-025-00398-6 [Download]
Browning, H. & Veit, W. (2025). Longtermism and animals. In H. Greaves, J. Barrett & D. Thorstad (eds). Essays on Longtermism. Oxford: Oxford University Press, pp. 449-462
https://doi.org/10.1093/9780191979972.003.0028 [Download]
Browning, H. & Veit, W. (2025). Self-awareness and personhood in non-human animals. In B.K. Keltz (ed). The Palgrave Handbook on the Problem of Animal Suffering in the Philosophy of Religion. Cham: Palgrave-Macmillan, pp. 475-498.
https://doi.org/10.1007/978-3-031-88233-3_22
Browning, H. & Veit, W. (2025). Best practice for farmed insect welfare: Lessons from Black Soldier Flies. CABI Animal Behaviour and Welfare Cases. abwcases20250014
https://doi.org/10.1079/abwcases.2025.0014. [Download preprint]
Veit, W., Browning, H., Garcia-Pelegrin, E., Davies, J.R., DuBois, J. & Clayton, N. (2025). Dimensions of corvid consciousness. Animal Cognition. 28:35
https://doi.org/10.1007/s10071-025-01949-y [Download]
Browning, H., Burn, C., Schnell, A., Crump, A. & Birch, J. (2025). Animal welfare risks from commercial practices involving cephalopod molluscs and decapod crustaceans. Animal Welfare 34(e24)
https://doi.org/10.1017/awf.2025.25 [Download]
Browning, H. & Veit, W. (2025). Animal welfare, agency, and animal-computer interaction. Animals. 15(2): 219
https://doi.org/10.3390/ani15020219 [Download]
Browning, H. & Veit, W. (2025). Studying introspection in animals and AIs. In F. Kammerer & K. Frankish (eds). The Landscape of Introspection: What Forms Could Introspective Systems Take? Exeter: Imprint Academic, pp. 52-63
https://www.imprint.co.uk/product/landscape/
Browning, H. & Veit, W. (2023). Positive wild animal welfare. Biology and Philosophy. 38(14): 1-19
https://doi.org/10.1007/s10539-023-09901-5 [Download preprint]
Browning, H. & Veit, W. (2023). Studying animal feelings: Integrating sentience research and welfare science. Journal of Consciousness Studies. 30(7-8): 196-222
https://doi.org/10.53765/20512201.30.7.196
Anomaly, J., Browning, H., Fleischman, D. & Veit, W. (2023). Flesh without blood: The public health benefits of lab-grown meat. Bioethical Enquiry. 21: 167-175
https://doi.org/10.1007/s11673-023-10254-7 [Download]
Veit, W. & Browning, H. (2023). Feminism and enhancement. In Edwards M.L. & Palermos, S.O. (eds). Feminist Philosophy and Emerging Technologies. New York: Routledge, pp. 37-55
https://doi.org/10.4324/9781003275992-4
Browning, H. & Veit, W. (2023). The welfare of brain organoids. Molecular Psychology. 2(4): 1-9
https://doi.org/10.12688/molpsychol.17523.1
Crump, A., Browning, H., Schnell, A., Burn, C., and Birch, J. (2022). Invertebrate sentience and sustainable seafood. Nature Food. 3: 884-886
https://doi.org/10.1038/s43016-022-00632-6 [View]
Crump, A., Browning, H., Schnell, A., Burn, C., and Birch, J. (2022). Animal sentience research: Synthesis and proposals. Animal Sentience. 32(31)
https://doi.org/10.51291/2377-7478.1770 [Download]
Crump, A., Browning, H., Schnell, A., Burn, C., and Birch, J. (2022) Sentience in decapod crustaceans: A general framework and review of the evidence. Animal Sentience. 32(1)
https://doi.org/10.51291/2377-7478.1691 [Download]
Browning, H. & Birch, J. (2022). Animal sentience. Philosophy Compass. e12822: 1-14
https://doi.org/10.1111/phc3.12822 [Download]
Browning, H. & Veit, W. (2022). The sentience shift in animal research. The New Bioethics. 28(4): 299-314
https://doi.org/10.1080/20502877.2022.2077681 [Download]
Browning, H. & Veit, W. (2022). The importance of end-of-life welfare. Animal Frontiers. 12(1): 8-15
https://doi.org/10.1093/af/vfab078 [Download]
Browning, H. & Veit, W. (2022). Why are we here? Evangelion and the desperate search for meaning in life. In Cotton, C. & Winters, A.M. (eds.) Neon Genesis Evangelion and Philosophy. Chicago: Carus Books, pp. 3-12
<Link to book> [Download preprint]
Birch, J., Burn, C., Schnell, A., Browning, H. & Crump, A. (2021). Review of the Evidence of Sentience in Cephalopod Molluscs and Decapod Crustaceans. London: Department for Environment, Food and Rural Affairs.
[Download]
Browning, H. & Veit, W. (2021). Freedom and animal welfare. Animals. 11(4): 1148
https://doi.org/10.3390/ani11041148 [Download]
Veit, W. & Browning, H. (2021). Developmental programming, evolution, and animal welfare: A case for evolutionary veterinary science. Journal of Applied Animal Welfare Science. 26(4): 552-564
https://doi.org/10.1080/10888705.2021.2014838 [Download]
Veit, W. & Browning, H. (2021). Phenomenology applied to animal health and suffering. In S. Ferrarello (Ed.), Phenomenology of Bioethics: Technoethics and Lived-Experience. Springer: pp. 73-88
https://doi.org/10.1007/978-3-030-65613-3_6 [Download preprint]
Veit, W. & Browning, H. (2021). Ethics of Mixed Martial Arts. In Holt, J. & Ramsay, M. (Ed.), The Philosophy of Mixed Martial Arts: Squaring the Octagon. London: Routledge, pp. 134-149.
https://doi.org/10.4324/9781003122395-12 [Download preprint]
Browning, H. & Veit, W. (2020). The Measurement Problem of Consciousness. Philosophical Topics. 48(1): 85-108.
https://doi.org/10.5840/philtopics20204815 [Download preprint]
Veit, W. & Browning, H. (2020). Perspectival pluralism for animal welfare. European Journal for Philosophy of Science. 11(9). https://doi.org/10.1007/s13194-020-00322-9 [Download]
Browning, H. & Veit, W. (2020). Is humane slaughter possible? Animals. 10(5): 799
https://doi.org/10.3390/ani10050799 [Download]
Browning, H. & Veit, W. (2020). Confined freedom and free confinement: The ethics of captivity in Life of Pi. In Á. T. Bogár and R. S. Szigethy (Eds.). Critical Insights: Life of Pi. Salem Press: 119-134.
https://doi.org/10.13140/RG.2.2.27969.38246/1 <Link to Book> [Download preprint]
Browning, H. & Maple, T. (2019). Developing a metric of usable space for zoo exhibits. Frontiers in Psychology. 10(791) https://doi.org/10.3389/fpsyg.2019.00791 [Download]
Browning-Jones, H. & Moro, L. (2006). A multi-sensory enrichment program for ring-tailed lemurs (Lemur catta) at Auckland Zoo, including a novel feeding device. In Proceedings of the 1st Australasian Regional Environmental Enrichment Conference (pp. 42-47).
https://doi.org/10.13140/RG.2.2.10908.92809 [Download]
Veit, W. & Browning, H. (forthcoming). Do animals understand death, and does it matter? In The Palgrave Handbook on the Philosophy of Death.
Browning, H. & Veit, W. (forthcoming). From animal to AI welfare. In Z. Radman (ed.). HumANimal: The “Animal Turn” Meets Artificial Intelligence. Palgrave MacMillan.
Browning, H. & Veit, W. (forthcoming). Does the welfare of wildlife matter? In A. Simmons (ed.). The Welfare of Wildlife.
Veit, W. & Browning, H. (forthcoming). Animal desire. In A. Gregory (ed.). The Routledge Handbook on the Philosophy of Desire.
Browning. H. & Powell, D. (2026). Navigating welfare contradictions in zoo-supported conservation activities. Zoo Biology: e70100
https://doi.org/10.1002/zoo.70100 [Download]
Webb, L.E., Bolhuis, J.E., Jurkovich, V., Ipema, A.F., Chincarini, M., Favreau-Peigné, A., Forkman, B., Clouard, C., Tuyttens, F., Browning, H., Menchetti, L., Bak Jensen, M., Elmaz, Ö ., Šárová, R., Głogowski, R., Duepjan, S., Celozzi, S., Veit, W., Tallet, C. (2026). Balancing the animal affective scales - Can the methodology of human affect balance be translated to other sentient species? Neuroscience & Biobehavioral Reviews. 106914
https://doi.org/10.1016/j.neubiorev.2026.106914
Elliott, V., Beausoleil, N., Browning, H., Díez-León, M., Goddard, P., Graham, M., Ramp, D., Sebo, J., Soryl, A. & Zemanova, M.A. (2026). The emerging field of wild animal welfare science. Trends in Ecology & Evolution. Online first.
https://doi.org/10.1016/j.tree.2026.06.003 [Download]
Keens Caballero, H., Browning, H., Lambton, S., Maye, D., & Roe, E. (2026). Resilience ontologies in veterinary science: How they shape the way we address resilience. Veterinary Sciences, 13(5): 471.
https://doi.org/10.3390/vetsci13050471 [Download]
Veit, W. & Browning, H. (2026). Two kinds of conceptual engineering. Croatian Journal of Philosophy. 26(76): 49-70.
https://doi.org/10.52685/cjp.26.76.4 [Download]
Schnell, A.K., Browning, H., Crump, A., Burn, C.C. & Birch, J. (2026). Sentience in cephalopod molluscs: An updated assessment. Biological Reviews. 101: 1311-1333
https://doi.org/10.1002/brv.70125
Ishida, S., Kagan, B.J., Kataoka, M., et al. (2026). Ethics and regulation of human brain organoid research: Recommendations from the Asia Pacific Neuroethics Working Group. Asian Bioethics Review. Online first.
https://doi.org/10.1007/s41649-025-00398-6 [Download]
Browning, H. & Veit, W. (2025). Longtermism and animals. In H. Greaves, J. Barrett & D. Thorstad (eds). Essays on Longtermism. Oxford: Oxford University Press, pp. 449-462
https://doi.org/10.1093/9780191979972.003.0028 [Download]
Browning, H. & Veit, W. (2025). Self-awareness and personhood in non-human animals. In B.K. Keltz (ed). The Palgrave Handbook on the Problem of Animal Suffering in the Philosophy of Religion. Cham: Palgrave-Macmillan, pp. 475-498.
https://doi.org/10.1007/978-3-031-88233-3_22
Browning, H. & Veit, W. (2025). Best practice for farmed insect welfare: Lessons from Black Soldier Flies. CABI Animal Behaviour and Welfare Cases. abwcases20250014
https://doi.org/10.1079/abwcases.2025.0014. [Download preprint]
Veit, W., Browning, H., Garcia-Pelegrin, E., Davies, J.R., DuBois, J. & Clayton, N. (2025). Dimensions of corvid consciousness. Animal Cognition. 28:35
https://doi.org/10.1007/s10071-025-01949-y [Download]
Browning, H., Burn, C., Schnell, A., Crump, A. & Birch, J. (2025). Animal welfare risks from commercial practices involving cephalopod molluscs and decapod crustaceans. Animal Welfare 34(e24)
https://doi.org/10.1017/awf.2025.25 [Download]
Browning, H. & Veit, W. (2025). Animal welfare, agency, and animal-computer interaction. Animals. 15(2): 219
https://doi.org/10.3390/ani15020219 [Download]
Browning, H. & Veit, W. (2025). Studying introspection in animals and AIs. In F. Kammerer & K. Frankish (eds). The Landscape of Introspection: What Forms Could Introspective Systems Take? Exeter: Imprint Academic, pp. 52-63
https://www.imprint.co.uk/product/landscape/
Browning, H. & Veit, W. (2023). Positive wild animal welfare. Biology and Philosophy. 38(14): 1-19
https://doi.org/10.1007/s10539-023-09901-5 [Download preprint]
Browning, H. & Veit, W. (2023). Studying animal feelings: Integrating sentience research and welfare science. Journal of Consciousness Studies. 30(7-8): 196-222
https://doi.org/10.53765/20512201.30.7.196
Anomaly, J., Browning, H., Fleischman, D. & Veit, W. (2023). Flesh without blood: The public health benefits of lab-grown meat. Bioethical Enquiry. 21: 167-175
https://doi.org/10.1007/s11673-023-10254-7 [Download]
Veit, W. & Browning, H. (2023). Feminism and enhancement. In Edwards M.L. & Palermos, S.O. (eds). Feminist Philosophy and Emerging Technologies. New York: Routledge, pp. 37-55
https://doi.org/10.4324/9781003275992-4
Browning, H. & Veit, W. (2023). The welfare of brain organoids. Molecular Psychology. 2(4): 1-9
https://doi.org/10.12688/molpsychol.17523.1
Crump, A., Browning, H., Schnell, A., Burn, C., and Birch, J. (2022). Invertebrate sentience and sustainable seafood. Nature Food. 3: 884-886
https://doi.org/10.1038/s43016-022-00632-6 [View]
Crump, A., Browning, H., Schnell, A., Burn, C., and Birch, J. (2022). Animal sentience research: Synthesis and proposals. Animal Sentience. 32(31)
https://doi.org/10.51291/2377-7478.1770 [Download]
Crump, A., Browning, H., Schnell, A., Burn, C., and Birch, J. (2022) Sentience in decapod crustaceans: A general framework and review of the evidence. Animal Sentience. 32(1)
https://doi.org/10.51291/2377-7478.1691 [Download]
Browning, H. & Birch, J. (2022). Animal sentience. Philosophy Compass. e12822: 1-14
https://doi.org/10.1111/phc3.12822 [Download]
Browning, H. & Veit, W. (2022). The sentience shift in animal research. The New Bioethics. 28(4): 299-314
https://doi.org/10.1080/20502877.2022.2077681 [Download]
Browning, H. & Veit, W. (2022). The importance of end-of-life welfare. Animal Frontiers. 12(1): 8-15
https://doi.org/10.1093/af/vfab078 [Download]
Browning, H. & Veit, W. (2022). Why are we here? Evangelion and the desperate search for meaning in life. In Cotton, C. & Winters, A.M. (eds.) Neon Genesis Evangelion and Philosophy. Chicago: Carus Books, pp. 3-12
<Link to book> [Download preprint]
Birch, J., Burn, C., Schnell, A., Browning, H. & Crump, A. (2021). Review of the Evidence of Sentience in Cephalopod Molluscs and Decapod Crustaceans. London: Department for Environment, Food and Rural Affairs.
[Download]
Browning, H. & Veit, W. (2021). Freedom and animal welfare. Animals. 11(4): 1148
https://doi.org/10.3390/ani11041148 [Download]
Veit, W. & Browning, H. (2021). Developmental programming, evolution, and animal welfare: A case for evolutionary veterinary science. Journal of Applied Animal Welfare Science. 26(4): 552-564
https://doi.org/10.1080/10888705.2021.2014838 [Download]
Veit, W. & Browning, H. (2021). Phenomenology applied to animal health and suffering. In S. Ferrarello (Ed.), Phenomenology of Bioethics: Technoethics and Lived-Experience. Springer: pp. 73-88
https://doi.org/10.1007/978-3-030-65613-3_6 [Download preprint]
Veit, W. & Browning, H. (2021). Ethics of Mixed Martial Arts. In Holt, J. & Ramsay, M. (Ed.), The Philosophy of Mixed Martial Arts: Squaring the Octagon. London: Routledge, pp. 134-149.
https://doi.org/10.4324/9781003122395-12 [Download preprint]
Browning, H. & Veit, W. (2020). The Measurement Problem of Consciousness. Philosophical Topics. 48(1): 85-108.
https://doi.org/10.5840/philtopics20204815 [Download preprint]
Veit, W. & Browning, H. (2020). Perspectival pluralism for animal welfare. European Journal for Philosophy of Science. 11(9). https://doi.org/10.1007/s13194-020-00322-9 [Download]
Browning, H. & Veit, W. (2020). Is humane slaughter possible? Animals. 10(5): 799
https://doi.org/10.3390/ani10050799 [Download]
Browning, H. & Veit, W. (2020). Confined freedom and free confinement: The ethics of captivity in Life of Pi. In Á. T. Bogár and R. S. Szigethy (Eds.). Critical Insights: Life of Pi. Salem Press: 119-134.
https://doi.org/10.13140/RG.2.2.27969.38246/1 <Link to Book> [Download preprint]
Browning, H. & Maple, T. (2019). Developing a metric of usable space for zoo exhibits. Frontiers in Psychology. 10(791) https://doi.org/10.3389/fpsyg.2019.00791 [Download]
Browning-Jones, H. & Moro, L. (2006). A multi-sensory enrichment program for ring-tailed lemurs (Lemur catta) at Auckland Zoo, including a novel feeding device. In Proceedings of the 1st Australasian Regional Environmental Enrichment Conference (pp. 42-47).
https://doi.org/10.13140/RG.2.2.10908.92809 [Download]
Coauthored Commentaries and Book Reviews
Browning, H. & Veit, W. (2026). Evolution, time, and the observer. Behavioral and Brain Sciences. 49: e170
https://doi.org/10.1017/S0140525X25102926
Dickel, L., Truong, S., Browning, H., Lynning Harfeld, J., & Rademann, A. (2026). Book review: Martha C. Nussbaum, Justice for Animals: Our Collective Responsibility, Simon & Schuster, New York (2020). Applied Animal Behaviour Science. 295: 106886
https://doi.org/10.1016/j.applanim.2025.106886
Browning, H. & Veit, W. (2025). Book review: Nigel Rothfels. Savages and beasts: the birth of the modern zoo (Revised Edition). 2025. Johns Hopkins University Press, Baltimore. History and Philosophy of the Life Sciences. 47(40): 1-4
https://doi.org/10.1007/s40656-025-00690-6 [View online]
Browning, H. & Veit, W. (2025). The neurodiversity model and medical model: Competitors or alternative perspectives? AJOB Neuroscience. 61(1): 32-34
https://doi.org/10.1080/21507740.2024.2437994
Jacquet, J. et al. (2024). Support US OCTOPUS Act to keep octopuses wild. Science. 385 (6710): 721-22
https://doi.org/10.1126/science.adr3813
Veit, W. & Browning, H. (2024). Meta-learning and the evolution of cognition. Behavioral and Brain Sciences. 47 (e167)
https://doi.org/10.1017/S0140525X24000177
Browning, H. & Veit, W. (2024). Animal welfare science, performance metrics, and proxy failure. Behavioral and Brain Sciences. 47 (e70)
https://doi.org/10.1017/S0140525X2300300X
Browning, H. & Veit, W. (2024). Book review: Saving Animals, Saving Ourselves. Ethics. 134 (3): 443-447
https://doi.org/10.1086/728630
Browning, H. & Veit, W. (2023). Studying introspection in animals and AIs. Journal of Consciousness Studies. 30 (9-10): 63-74
https://doi.org/10.53765/20512201.30.9.063
Jacquet, J., Franks, B., Browning, H., Donoso, A., Hayek, M., Hessler, K. & Sellars, L. (2023). Away from an offshore aquaculture industry in the United States. Marine Policy. 160: 105968
https://doi.org/10.1016/j.marpol.2023.105986
Browning, H. & Veit, W. (2023). Moral Awareness and Animal Welfare (book review). Animal Welfare. 32 (e39)
https://doi.org/10.1017/awf.2023.33
Veit, W. & Browning, H. (2023). Neural networks, AI, and the goals of modeling. Behavioral and Brain Sciences. 46 (e411)
https://doi.org/10.1017/S0140525X23001681
Veit, W. & Browning, H. (2023). Evolutionary mismatch and anomalies in the memory system. Behavioral and Brain Sciences. 46 (e381)
https://doi.org/10.1017/S0140525X23000183
Veit, W. & Browning, H. (2023). Puritanical morality and the scaffolded evolution of self-control. Behavioral and Brain Sciences. 46 (e319)
https://doi.org/10.1017/S0140525X23000535
Veit, W. & Browning, H. (2023). The design space of human communication and the nonevolution of ideography. Behavioral and Brain Sciences. 46 (e255)
https://doi.org/10.1017/S0140525X23000754
Veit, W. & Browning, H. (2023). Polygenic scores and social science. Behavioral and Brain Sciences. 46 (e229)
https://doi.org/10.1017/S0140525X22002345
Veit, W. & Browning, H. (2023). Hominin life history, pathological complexity, and the evolution of anxiety. Behavioral and Brain Sciences. 46 (e79)
https://doi.org/10.1017/S0140525X22001923 [Download preprint]
Browning, H. & Veit, W. (2023). Regulating possibly sentient human cerebral organoids. AJOB Neuroscience. 14(2): 197-199
https://doi.org/10.1080/21507740.2023.2188293
Veit, W. & Browning, H. (2023). Defending sentientism. AJOB Neuroscience. 14(2): 168-170
https://doi.org/10.1080/21507740.2023.2188292
Veit, W. & Browning, H. (2023). Social robots and the intentional stance. Behavioral and Brain Sciences. 46 (e47)
https://doi.org/10.1017/S0140525X22001595 [Download preprint]
Browning, H. & Veit, W. (2023). Optimism about measuring animal feelings. Asian Bioethics Review: 1-5
https://doi.org/10.1007/s41649-023-00244-7
Veit, W. & Browning, H. (2023). The scaffolded evolution of human communication. Behavioral and Brain Sciences. 46 (e17)
https://doi.org/10.1017/S0140525X2200070X [Download preprint]
Veit W. & Browning, H. (2022). Darwinian and autopoietic views of the organism. Constructivist Foundations. 18(1): 103–105.
https://constructivist.info/18/1/103
Veit, W. & Browning, H. (2022). On the evolutionary origins of the bifocal stance. Behavioral and Brain Sciences. 45 (e270)
https://doi.org/10.1017/S0140525X22001273 [Download preprint]
Veit, W. & Browning, H. (2022). Life, mind, agency: Why Markov blankets fail the test of evolution. Behavioral and Brain Sciences. 45 (e214)
https://doi.org/10.1017/S0140525X22000115 [Download preprint]
Browning, H. & Veit, W. (2022). Autism and the preference for imaginary worlds. Behavioral and Brain Sciences. 45 (e279)
https://doi.org/10.1017/S0140525X21002211 [Download preprint]
Veit, W. & Browning, H. (2022). Pathological complexity and the evolution of sex differences. Behavioral and Brain Sciences. 45(149): 57-58
https://www.doi.org/10.1017/S0140525X22000498. [Download preprint]
Browning, H. & Veit, W. (2022). More than zombies: Considering the animal subject in de-extinction. Ethics, Policy & Environment. 25(2): 121-124
https://doi.org/10.1080/21550085.2022.2071552 [Download preprint]
Birch, J., Broom, D.M., Browning, H., Crump, A., Ginsburg, S., Halina, M., Harrison, D., Jablonka, E., Lee, A.Y., Kammerer, F. and Klein, C. (2022). How should we study animal consciousness scientifically? Journal of Consciousness Studies. 29(3-4): 8-28
https://doi.org/10.53765/20512201.29.3.008 [Download]
Browning, H. & Veit, W. (2021). Evolutionary biology meets consciousness: Essay review of Simona Ginsburg and Eva Jablonka’s The Evolution of the Sensitive Soul. Biology and Philosophy. 36(1): 5
https://doi.org/10.1007/s10539-021-09781-7 [Download]
Browning, H. & Veit, W. (2021). Does utilitarianism need a rethink? Review of Louis Narens and Brian Skyrms’ The Pursuit of Happiness. Journal of Economic Methodology. 29(3): 256-261
https://doi.org/10.1080/1350178X.2021.1985755 [Download preprint]
Veit, W. & Browning, H. (2021). Extending animal welfare science to include wild animals. Animal Sentience. 31(1).
https://doi.org/10.51291/2377-7478.1675 [Download]
Birch, J. & Browning, H. (2021). Neural organoids and the precautionary principle. The American Journal of Bioethics. 21(1): 56-58.
https://doi.org/10.1080/15265161.2020.1845858 [Download preprint]
Browning, H. & Veit, W. (2021). On the relevance of experimental philosophy to neuroethics. AJOB Neuroscience. 13(1): 55-57
https://doi.org/10.1080/21507740.2021.2001080 [Download preprint]
Veit, W. & Browning, H. (2021). Has the socio-political role of neuroethics been neglected? AJOB Neuroscience. 13(1): 23-25
https://doi.org/10.1080/21507740.2021.2001079 [Download preprint]
Browning, H. & Veit, W. (2021). Utilitarian lessons from the COVID-19 pandemic for non-pandemic diseases. American Journal of Bioethics. 21(12): 39-42
https://doi.org/10.1080/15265161.2021.1991035 [Download preprint]
Veit, W., Earp, B.D., Browning, H., & Savulescu, J. (2021). Evaluating trade-offs between autonomy and wellbeing in supported decision making. American Journal of Bioethics. 21(11): 21-24
https://doi.org/10.1080/15265161.2021.1980134 [Download preprint]
Veit, W. & Browning, H. (2020). Why socio-political beliefs trump individual morality: An evolutionary perspective. AJOB Neuroscience. 11(4): 290-292.
https://doi.org/10.1080/21507740.2020.1830879
Browning, H. & Veit, W. (2020). Improving invertebrate welfare. Animal Sentience. 29(4)
http://doi.org/10.51291/2377-7478.1585 [Download]
Browning, H. & Veit, W. (2026). Evolution, time, and the observer. Behavioral and Brain Sciences. 49: e170
https://doi.org/10.1017/S0140525X25102926
Dickel, L., Truong, S., Browning, H., Lynning Harfeld, J., & Rademann, A. (2026). Book review: Martha C. Nussbaum, Justice for Animals: Our Collective Responsibility, Simon & Schuster, New York (2020). Applied Animal Behaviour Science. 295: 106886
https://doi.org/10.1016/j.applanim.2025.106886
Browning, H. & Veit, W. (2025). Book review: Nigel Rothfels. Savages and beasts: the birth of the modern zoo (Revised Edition). 2025. Johns Hopkins University Press, Baltimore. History and Philosophy of the Life Sciences. 47(40): 1-4
https://doi.org/10.1007/s40656-025-00690-6 [View online]
Browning, H. & Veit, W. (2025). The neurodiversity model and medical model: Competitors or alternative perspectives? AJOB Neuroscience. 61(1): 32-34
https://doi.org/10.1080/21507740.2024.2437994
Jacquet, J. et al. (2024). Support US OCTOPUS Act to keep octopuses wild. Science. 385 (6710): 721-22
https://doi.org/10.1126/science.adr3813
Veit, W. & Browning, H. (2024). Meta-learning and the evolution of cognition. Behavioral and Brain Sciences. 47 (e167)
https://doi.org/10.1017/S0140525X24000177
Browning, H. & Veit, W. (2024). Animal welfare science, performance metrics, and proxy failure. Behavioral and Brain Sciences. 47 (e70)
https://doi.org/10.1017/S0140525X2300300X
Browning, H. & Veit, W. (2024). Book review: Saving Animals, Saving Ourselves. Ethics. 134 (3): 443-447
https://doi.org/10.1086/728630
Browning, H. & Veit, W. (2023). Studying introspection in animals and AIs. Journal of Consciousness Studies. 30 (9-10): 63-74
https://doi.org/10.53765/20512201.30.9.063
Jacquet, J., Franks, B., Browning, H., Donoso, A., Hayek, M., Hessler, K. & Sellars, L. (2023). Away from an offshore aquaculture industry in the United States. Marine Policy. 160: 105968
https://doi.org/10.1016/j.marpol.2023.105986
Browning, H. & Veit, W. (2023). Moral Awareness and Animal Welfare (book review). Animal Welfare. 32 (e39)
https://doi.org/10.1017/awf.2023.33
Veit, W. & Browning, H. (2023). Neural networks, AI, and the goals of modeling. Behavioral and Brain Sciences. 46 (e411)
https://doi.org/10.1017/S0140525X23001681
Veit, W. & Browning, H. (2023). Evolutionary mismatch and anomalies in the memory system. Behavioral and Brain Sciences. 46 (e381)
https://doi.org/10.1017/S0140525X23000183
Veit, W. & Browning, H. (2023). Puritanical morality and the scaffolded evolution of self-control. Behavioral and Brain Sciences. 46 (e319)
https://doi.org/10.1017/S0140525X23000535
Veit, W. & Browning, H. (2023). The design space of human communication and the nonevolution of ideography. Behavioral and Brain Sciences. 46 (e255)
https://doi.org/10.1017/S0140525X23000754
Veit, W. & Browning, H. (2023). Polygenic scores and social science. Behavioral and Brain Sciences. 46 (e229)
https://doi.org/10.1017/S0140525X22002345
Veit, W. & Browning, H. (2023). Hominin life history, pathological complexity, and the evolution of anxiety. Behavioral and Brain Sciences. 46 (e79)
https://doi.org/10.1017/S0140525X22001923 [Download preprint]
Browning, H. & Veit, W. (2023). Regulating possibly sentient human cerebral organoids. AJOB Neuroscience. 14(2): 197-199
https://doi.org/10.1080/21507740.2023.2188293
Veit, W. & Browning, H. (2023). Defending sentientism. AJOB Neuroscience. 14(2): 168-170
https://doi.org/10.1080/21507740.2023.2188292
Veit, W. & Browning, H. (2023). Social robots and the intentional stance. Behavioral and Brain Sciences. 46 (e47)
https://doi.org/10.1017/S0140525X22001595 [Download preprint]
Browning, H. & Veit, W. (2023). Optimism about measuring animal feelings. Asian Bioethics Review: 1-5
https://doi.org/10.1007/s41649-023-00244-7
Veit, W. & Browning, H. (2023). The scaffolded evolution of human communication. Behavioral and Brain Sciences. 46 (e17)
https://doi.org/10.1017/S0140525X2200070X [Download preprint]
Veit W. & Browning, H. (2022). Darwinian and autopoietic views of the organism. Constructivist Foundations. 18(1): 103–105.
https://constructivist.info/18/1/103
Veit, W. & Browning, H. (2022). On the evolutionary origins of the bifocal stance. Behavioral and Brain Sciences. 45 (e270)
https://doi.org/10.1017/S0140525X22001273 [Download preprint]
Veit, W. & Browning, H. (2022). Life, mind, agency: Why Markov blankets fail the test of evolution. Behavioral and Brain Sciences. 45 (e214)
https://doi.org/10.1017/S0140525X22000115 [Download preprint]
Browning, H. & Veit, W. (2022). Autism and the preference for imaginary worlds. Behavioral and Brain Sciences. 45 (e279)
https://doi.org/10.1017/S0140525X21002211 [Download preprint]
Veit, W. & Browning, H. (2022). Pathological complexity and the evolution of sex differences. Behavioral and Brain Sciences. 45(149): 57-58
https://www.doi.org/10.1017/S0140525X22000498. [Download preprint]
Browning, H. & Veit, W. (2022). More than zombies: Considering the animal subject in de-extinction. Ethics, Policy & Environment. 25(2): 121-124
https://doi.org/10.1080/21550085.2022.2071552 [Download preprint]
Birch, J., Broom, D.M., Browning, H., Crump, A., Ginsburg, S., Halina, M., Harrison, D., Jablonka, E., Lee, A.Y., Kammerer, F. and Klein, C. (2022). How should we study animal consciousness scientifically? Journal of Consciousness Studies. 29(3-4): 8-28
https://doi.org/10.53765/20512201.29.3.008 [Download]
Browning, H. & Veit, W. (2021). Evolutionary biology meets consciousness: Essay review of Simona Ginsburg and Eva Jablonka’s The Evolution of the Sensitive Soul. Biology and Philosophy. 36(1): 5
https://doi.org/10.1007/s10539-021-09781-7 [Download]
Browning, H. & Veit, W. (2021). Does utilitarianism need a rethink? Review of Louis Narens and Brian Skyrms’ The Pursuit of Happiness. Journal of Economic Methodology. 29(3): 256-261
https://doi.org/10.1080/1350178X.2021.1985755 [Download preprint]
Veit, W. & Browning, H. (2021). Extending animal welfare science to include wild animals. Animal Sentience. 31(1).
https://doi.org/10.51291/2377-7478.1675 [Download]
Birch, J. & Browning, H. (2021). Neural organoids and the precautionary principle. The American Journal of Bioethics. 21(1): 56-58.
https://doi.org/10.1080/15265161.2020.1845858 [Download preprint]
Browning, H. & Veit, W. (2021). On the relevance of experimental philosophy to neuroethics. AJOB Neuroscience. 13(1): 55-57
https://doi.org/10.1080/21507740.2021.2001080 [Download preprint]
Veit, W. & Browning, H. (2021). Has the socio-political role of neuroethics been neglected? AJOB Neuroscience. 13(1): 23-25
https://doi.org/10.1080/21507740.2021.2001079 [Download preprint]
Browning, H. & Veit, W. (2021). Utilitarian lessons from the COVID-19 pandemic for non-pandemic diseases. American Journal of Bioethics. 21(12): 39-42
https://doi.org/10.1080/15265161.2021.1991035 [Download preprint]
Veit, W., Earp, B.D., Browning, H., & Savulescu, J. (2021). Evaluating trade-offs between autonomy and wellbeing in supported decision making. American Journal of Bioethics. 21(11): 21-24
https://doi.org/10.1080/15265161.2021.1980134 [Download preprint]
Veit, W. & Browning, H. (2020). Why socio-political beliefs trump individual morality: An evolutionary perspective. AJOB Neuroscience. 11(4): 290-292.
https://doi.org/10.1080/21507740.2020.1830879
Browning, H. & Veit, W. (2020). Improving invertebrate welfare. Animal Sentience. 29(4)
http://doi.org/10.51291/2377-7478.1585 [Download]
Public Engagement
Browning, H. (2025). How citizens’ assemblies could improve animal welfare. The Conversation.
https://theconversation.com/how-citizens-assemblies-could-improve-animal-welfare-259755
Veit, W. & Browning, H. (2025). What’s it like being a raven or a crow? The Conversation.
https://theconversation.com/whats-it-like-being-a-raven-or-a-crow-257171
Browning, H. (2025). Science brought the dire wolf back – but who’s thinking about its welfare? LBC Opinion.
https://www.lbc.co.uk/opinion/views/science-dire-wolf-animal-welfare/
Browning, H. (2023). Do the potential harms of de-extinction outweigh the benefits? CQ Researcher.
https://cqpress.sagepub.com/cqresearcher/report/cqpromo/yya2V8/reversing-extinction-cqresrre20230901
Browning, H. & Veit, W. (2023). What is life like as a wild animal? Probably nicer than you think. The Conversation.
https://theconversation.com/what-is-life-like-as-a-wild-animal-probably-nicer-than-you-think-205322
Browning, H., Schnell, A. & Birch, J. (2022). Octopus farms raise huge animal welfare concerns - and they’re unsustainable too. The Conversation.
https://theconversation.com/octopus-farms-raise-huge-animal-welfare-concerns-and-theyre-unsustainable-too-179134
Browning, H. (2025). How citizens’ assemblies could improve animal welfare. The Conversation.
https://theconversation.com/how-citizens-assemblies-could-improve-animal-welfare-259755
Veit, W. & Browning, H. (2025). What’s it like being a raven or a crow? The Conversation.
https://theconversation.com/whats-it-like-being-a-raven-or-a-crow-257171
Browning, H. (2025). Science brought the dire wolf back – but who’s thinking about its welfare? LBC Opinion.
https://www.lbc.co.uk/opinion/views/science-dire-wolf-animal-welfare/
Browning, H. (2023). Do the potential harms of de-extinction outweigh the benefits? CQ Researcher.
https://cqpress.sagepub.com/cqresearcher/report/cqpromo/yya2V8/reversing-extinction-cqresrre20230901
Browning, H. & Veit, W. (2023). What is life like as a wild animal? Probably nicer than you think. The Conversation.
https://theconversation.com/what-is-life-like-as-a-wild-animal-probably-nicer-than-you-think-205322
Browning, H., Schnell, A. & Birch, J. (2022). Octopus farms raise huge animal welfare concerns - and they’re unsustainable too. The Conversation.
https://theconversation.com/octopus-farms-raise-huge-animal-welfare-concerns-and-theyre-unsustainable-too-179134
